BS: Biology, State University of Utrecht, the Netherlands, 1983.
MS: Medical Biology, State University of Utrecht, the Netherlands, 1987.
Teaching degree: Biology, State University of Utrecht, Netherlands, 1989.
PhD: Hematology, Erasmus University Rotterdam, the Netherlands, 1995.
Postdoctoral Fellowship: Hematology, Howard Hughes Medical Institute, Indiana University, Indianapolis, IN, 1994-1998.